www.sciencedirect.com/topics/agricultural-and-biological-sciences/dune-soils
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Humus accumulation causes soil acidity and dunes thus become more acidic as they age.
n Studland, Dorset, the early invasion of Calluna is largely responsible for a very rapid increase in organic matter.
In dune depressions, where water is not a limiting factor for plant establishment, the accumulation of organic matter is faster.
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